↑ Staeck, W.; Schindler, I. (2015). “Description of a new Heros species (Teleostei, Cichlidae) from the Rio Orinoco drainage and notes on Heros severus Heckel, 1840”. Bulletin of Fish Biology. 15 (1–2): 121–136.
1 2 Kullander, S.O., 2003. Cichlidae (Cichlids). p. 605-654. A R.E. Reis, S.O. Kullander y C.J. Ferraris, Jr. (eds.) Checklist of the Freshwater Fishes of South and Central America. Porto Alegre: EDIPUCRS, Brasil.
